Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males

Seksuaalivalinta aiheuttaa yksilöiden välille kilpailua parittelukumppaneista, minkä johdosta osa yksilöistä lisääntyy toisia enemmän. Elinkiertoteorian mukaan yksilöt eivät niukkojen resurssien vuoksi pysty tehokkaasti kohdentamaan energiaa samanaikaisesti useisiin toimintoihin, kuten kasvuun, selv...

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Main Author: Nieminen, Eini
Other Authors: Matemaattis-luonnontieteellinen tiedekunta, Faculty of Sciences, Bio- ja ympäristötieteiden laitos, Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylän yliopisto
Format: Master's thesis
Language:eng
Published: 2014
Subjects:
Online Access: https://jyx.jyu.fi/handle/123456789/44793
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author Nieminen, Eini
author2 Matemaattis-luonnontieteellinen tiedekunta Faculty of Sciences Bio- ja ympäristötieteiden laitos Department of Biological and Environmental Science University of Jyväskylä Jyväskylän yliopisto
author_facet Nieminen, Eini Matemaattis-luonnontieteellinen tiedekunta Faculty of Sciences Bio- ja ympäristötieteiden laitos Department of Biological and Environmental Science University of Jyväskylä Jyväskylän yliopisto Nieminen, Eini Matemaattis-luonnontieteellinen tiedekunta Faculty of Sciences Bio- ja ympäristötieteiden laitos Department of Biological and Environmental Science University of Jyväskylä Jyväskylän yliopisto
author_sort Nieminen, Eini
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description Seksuaalivalinta aiheuttaa yksilöiden välille kilpailua parittelukumppaneista, minkä johdosta osa yksilöistä lisääntyy toisia enemmän. Elinkiertoteorian mukaan yksilöt eivät niukkojen resurssien vuoksi pysty tehokkaasti kohdentamaan energiaa samanaikaisesti useisiin toimintoihin, kuten kasvuun, selviytymiseen ja lisääntymiseen. Ikä, suuri koko ja näyttävät seksuaaliornamentit ovat seksuaalivalinnassa menestyvän yksilön merkkejä, sillä tällaisilla yksilöillä riittää energiaa elintärkeiden toimintojen lisäksi myös lisääntymiseen. Tavallisesti ne menestyvätkin parhaiten kilpailussa parittelukumppaneista. Nuoret tai huonokuntoiset yksilöt voivat kuitenkin turvautua vaihtoehtoisiin lisääntymisstrategioihin välttääkseen suoraa kilpailua dominoivien yksilöiden kanssa ja parantaakseen lisääntymismenestystään. Tässä tutkimuksessa perehdyttiin nuorten teerikukkojen (Tetrao tetrix) soidinvireyden ajoittamiseen vertaamalla 1-vuotiaiden ja aikuisten kukkojen soidinkäyttäytymistä eli läsnäoloa ja tappeluaktiivisuutta, sekä morfologisten ominaisuuksien eli ruumiinpainon ja pyrstön pituuden vaikutusta soidinkäyttäytymiseen. Mittava pitkäaikaisaineisto kerättiin vuosina 2003–2013 kolmelta keskisuomalaiselta soitimelta. Aineisto analysoitiin lineaarisia sekamalleja (GLMM) hyväksikäyttäen. Eniten soitimella läsnä olleiden nuorten kukkojen havaittiin ajoittavan soidinvireytensä aikuisia kukkoja myöhemmin. Tätä vaihtoehtoista lisääntymisstrategiaa noudattavat nuoret kukot olivat paitsi ahkerasti läsnä soitimella, myös aktiivisia tappelijoita ja kooltaan suurempia kuin muut 1-vuotiaat kukot. Todennäköisesti nämä nuoret kukot pystyivät panostamaan soidinkauteen huomattavasti ikätovereitaan enemmän, mutta kuitenkin aikuisia kukkoja vähemmän, minkä johdosta ne ehkä hyötyivät myöhemmästä soidinvireyden ajoituksesta eniten. Nuorten kukkojen vaihtoehtoinen lisääntymisstrategia voi asettaa lisää valintapaineita myös aikuisille kukoille hidastamalla kilpailun vähenemistä soidinkauden lopulla. Tämä tutkimus osoittaakin epäsuorasti, että niin sanotulla kestävyyskilpailulla (”endurance rivalry”) voi olla sijansa teeren seksuaalivalinnassa. Sexual selection generates competition over mates producing differences in reproductive success between individuals. The life history theory predicts that because of scarce resources individuals face the problem in allocating energy to different functions such as growth, survival, and reproduction. As a consequence, older age, large body size, and impressive sexual ornaments are the common characters of dominant individuals, who are often superior in competition over mates thus enjoying the highest reproductive success. Nevertheless, young or unfit individuals may have alternative mating strategies which assist them to avoid competition with superior individuals and increase their reproductive success. In this study timing of reproductive effort as a potentially alternative mating strategy of young lekking black grouse (Tetrao tetrix) males was investigated. The behavioural (lek attendance and fighting rate) features of yearling and adult males were compared, and the effect of morphological (body mass and tail length) traits on the behaviour were studied. The large long-term dataset was gathered in years 2003–2013 from three study sites, and analysed by generalized linear mixed effects models (GLMM). Yearling males, who were the most present on the lek arena, scheduled their reproductive effort later in the lekking season compared to adult males. The yearling males showing this alternative mating strategy proved to be of a high quality: compared to other yearlings, they were diligently present on the lek arena, fought actively and had larger body mass. Probably they were able to invest substantially more to the lekking season than most of the yearlings, but less than adults, and therefore profited from the alternative mating strategy. Later timing of effort in yearling males may also set more selection pressure for adult males by delaying the decrease of intensive competition in the end of the lekking season. Hence, this study supports indirectly the idea that endurance rivalry may operate on black grouse leks.
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spellingShingle Nieminen, Eini Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males energy allocation lek mate competition reproductive strategy selection Ekologia ja evoluutiobiologia Ecology and evolutionary biology 4011 teeri soidin lisääntyminen sukupuolivalinta
title Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
title_full Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
title_fullStr Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
title_full_unstemmed Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
title_short Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
title_sort timing of reproductive effort as an alternative mating strategy in young black grouse tetrao tetrix males
title_txtP Timing of reproductive effort as an alternative mating strategy in young black grouse (Tetrao tetrix) males
topic energy allocation lek mate competition reproductive strategy selection Ekologia ja evoluutiobiologia Ecology and evolutionary biology 4011 teeri soidin lisääntyminen sukupuolivalinta
topic_facet 4011 Ecology and evolutionary biology Ekologia ja evoluutiobiologia energy allocation lek lisääntyminen mate competition reproductive strategy selection soidin sukupuolivalinta teeri
url https://jyx.jyu.fi/handle/123456789/44793 http://www.urn.fi/URN:NBN:fi:jyu-201412023413
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